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You might encounter this problem when you're first learning ZBrush and just getting used to the interface. I've decided that I want to pose my model using the Transpose tool. So I have my demo soldier model loaded in the canvas and I know to activate the Transpose tool I just need to click the Move, Scale or Rotate buttons. As you may anticipate when I do this I'm going to encounter a problem. So I click on Move and instead of the Transpose handle I get this strange looking tool. This is the Transform Gyro.
I can rotate the model and move it around. You can see he's actually interacting with the back of the canvas there. But I don't have my Transpose handles. The reason this occurred is I forgot that I need to be in Edit mode to make the Transpose tool active. So I'm going to switch back to Draw, turn on Edit, and I can see this border's here indicating Edit mode and my button is on. Now when I click Move I have my Transpose tool and I'm ready to start working with it.
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